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Conversely, in the jaguar and also the jaguarundi Puma yagouaroundi, melanism is due to the expression of a dominant mutant allele instead. Bin Laden was born the grandson of Mohammed bin Awad bin Laden, the founder of one of the wealthiest non- royal families in Saudi Arabia, and the son of Osama bin Laden, whose partisan activities shaped his childhood. Bin Laden accompanied his father on his exile to Sudan from 1991 to 1996, and then to Afghanistan after that.